Basic M2M Self Reference Through Question

I apologize if this is basic and I am just missing something in the django docs.  The Django ORM allows you to do a self reference through another table.  I understand everything about this process EXCEPT how Django determines which of the foreign keys to use for each side of the M2M relationship.

class MyUser(...):

    connections = models.ManyToManyField('self', through=Relationship, symmetrical=False)

class Relationship(...):

    follower = models.ForeignKey(MyUser)

    following = models.ForeignKey(MyUser)

So in the psuedocode example above how does Django ORM know which of the 2 foreign key fields to connect to?  Does it just assume the first field is the one to use?  Thanks for any help on this.

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